Davies returns from injury to reunite Wales’ most capped centre pairing as he lines up alongside Jamie Roberts (Racing Metro).
The only change in pack from the victory againstFrance sees Jones return in the second-row after missing the Millennium Stadium clash with a foot infection. He partners Luke Charteris (Perpignan) behind a front-row of Gethin Jenkins (Cardiff Blues), Richard Hibbard and Adam Jones (both Ospreys).
The back-row remains unchanged with captain Sam Warburton (Cardiff Blues) featuring alongside Dan Lydiate (Racing Metro) and Taulupe Faletau (NewportGwent Dragons).
Rhys Webb (Ospreys) and Rhys Priestland (Scarlets) continue their partnership at half-back.
There is one positional change in the back-line with George North (Northampton Saints) returning to the wing from centre and he lines up alongside Cardiff Blues pair Alex Cuthbert and Leigh Halfpenny.
“It’s great to have both Jon and Alun Wyn back and available and they come into a very experienced side,” said Gatland.
“Jon came through the match well at the weekend and he is ready to go. We were happy with the response and performance against France but we know we will have to raise it another level on Sunday.
“We are really looking forward to going up to Twickenham, England have performed well so far and its set up to be a great game with all to play for.”
On the bench Ken Owens, Rhodri Jones (both Scarlets) and Paul James (Bath Rugby) are named as the front-row replacements and arejoined by Scarlets second-row Jake Ball and Ospreys flanker Justin Tipuric. Mike Phillips (Racing Metro), Dan Biggar (Ospreys) and Liam Williams (Scarlets) provide cover for the backs.
WALES SQUAD TO PLAY ENGLAND:
LeighHalfpenny (Cardiff Blues), Alex Cuthbert (Cardiff Blues), Jonathan Davies (Scarlets), Jamie Roberts (Racing Metro), George North (Northampton Saints), Rhys Priestland (Scarlets), Rhys Webb (Ospreys), Gethin Jenkins (Cardiff Blues), Richard Hibbard (Ospreys), Adam Jones (Ospreys), Luke Charteris (Perpignan), Alun Wyn Jones (Ospreys), Dan Lydiate (Racing Metro), Sam Warburton (CAPT – Cardiff Blues), Taulupe Faletau (Newport Gwent Dragons).
Replacements: Ken Owens (Scarlets), Paul James (Bath Rugby), Rhodri Jones (Scarlets), Jake Ball (Scarlets), Justin Tipuric (Ospreys), Mike Phillips (Racing Metro), Dan Biggar (Ospreys), Liam Williams (Scarlets).